Canada seems like a good all-rounder, with a healthy commercial industry and a strong independent community (thanks to its National Film Board). The USA leads the world when it comes to technological sophistication, but unfortunately its less mainstream animation never seems to have had the support of an organisation comparable to the National Film Board of Canada or the British Channel 4.
Japan has its massive industry, and its indies seem to be getting more attention in the international scene lately, what with two Japanese shorts getting Oscar nods in the past few years (and one winning).